基金Supported by the Agriculture Science Technology Achievement Transformation Fund （No. 2014GB2C22015）, the Public Projects of Zhejiang Province （Nos. 2013C32037, 2013C31032）, the Zhejiang Major Special Program of Breeding （No. 2012C 12907-3）, the Ningbo Innovation and Entrepreneurship Project （No. 2014C92011）, and the Zhejiang Provincial Oceanic and Fishery Bureau Project, Zhoushan Science and Technology Bureau Project （No. 2013C41007）.ACKNOWLEDGMENT The funders had no role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript. The authors thank the laboratory members for the technical assistance, and are also grateful to the Marine Fishery Research Institute of Zhejiang Province for providing the swimming crabs.